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Many of the articles are reprinted from "The Dredgings" - Newsletter of the Pacific Northwest Shell Club

Newly Added
Other Mollusks and Invertebrates
Field Trips/Shelling Adventures
     (file sizes over 500kb are noted)

Newly Added

Giant Tridacna Clam - A Personal Perspective  (pdf)

How drift bottles were used to map Northern Pacific ocean currents in the 1950's (pdf)

Wasting-like lesions occurring on California Sea Cucumbers  (pdf)

Watching for Cepaea nemoralis to emerge from hibernation adds fun to late winter walks with my dog  (pdf)

Freshwater mollusc declines, local extinctions and introductions noted in five northern California streams  (pdf)

Correlating personal weather station data with Allogona townsendiana activity observations at Crystal Springs
Park, Tukwila, Washington  (pdf)

Mollusk Notes:  Neoteny in the Carinate Dove Shell, Alia carinata  (pdf)

Our Common Leopard Dorid Nudibranch has been separated into two species  (pdf)


A new species of deep-water bivalve, Ascetoaxinus quatsinoensis, from off the coast of Vancouver Island, B.C.  (pdf)

The Angelic Imposter. The False angel wing clam Petricolaria pholadiformis (Lamarck, 1818),
of Willapa Bay, Washington  (pdf)

Commarginal Bittersweet - A New Clam Species to the B.C. Fauna  (pdf)

Discovery and collection of the Varnished Mussel, Vilasina vernicosa -
      A southern range extension and a new Canadian record.  (pdf)

Effects of Ocean Acidification on Early Developmental Stages of the Pacific Oyster (Crassostrea gigas)
in an Aquaculture Setting  (pdf 10.5Mb)

Habitat use by juvenile mollusc species:  Nucella ostrina, Littorina scutulata and Mytilus trossulus  (pdf)

Harvesting Wrinkled Pen Shells, Pinna rugosa, in Baja California, Mexico (pdf)

Identifying our Large Intertidal Veneridae  (pdf)

In Search of the Duck-Bill Piddocks Netastoma japonicum and N. rostratum  (pdf)

Interior comparison of the shells of Intertidal Veneridae from the Pacific Northwest  (pdf)

Littlenecks for Food, Thought, and Enjoyment  (pdf)

Modiolus rectus Exposed  (pdf)

Modiolus rectus, The Mussel with a Mighty Byssus  (pdf)

Molluscs in Pholad Holes  (pdf)

A Mussel Attached to a Crab Inside a Clam  (pdf)

Native Clams Facilitate Invasive Species in an Eelgrass Bed  (pdf)

New Distribution Records & Habitat Information for TAYLOR'S DWARF-MUSSEL, 'Musculus' taylori (Dall, 1897)  (pdf)

New Species of Deepsea Bivalve Named for David McKay  (pdf)

Ocean acidification and its effect on the aquaculture industry  (pdf)

Pearl Found in Solen sicarius  (pdf)

Penitella Piddocks in Peat and Petrified Wood  (pdf)

[Paphia] Protothaca restorationensis Frizzell, 1930 - Series of 3 articles  (pdf)

Range Extension for Saxidomus nuttallii Conrad, 1837  (pdf)

A strange occurrence happening to siphons of Zirfaea pilsbryi in Puget Sound  (pdf)

A Third Tresus Found in Pacific Northwest  (pdf)


An Exciting new Discovery:  the Lightly-sculptured Odostome snail, Evalea tenuisculpta (Carpenter, 1864) feeding
on the siphon tips of the Fat Gaper, Tresus capax (Gould, 1850) in Vancouver Harbour, British Columbia  (pdf)

The Behavioral Ecology and Territioriality of the Owl Limpet, Lottia gigantea
     PhD. dissertation of Stephanie Lynn Schroeder Ph.D.  (pdf 2.95M)

The best of both worlds:  A clever intertidal snail feeds on land and cools off in the sea  (pdf)

Capulus ungaricus and friends  (pdf)

Collecting Lottia alveus [parallela] (Conrad, 1831) in Boundary Bay, British Columbia  (pdf)

The Common Periwinkle, Littorina littorea, in British Columbia  (pdf)

Crepidula convexa Say, 1822 discovered in Willapa Bay, WA  (pdf)

Crepidula onyx (G.B. Sowerby I, 1824) collected in Puget Sound and Pacific Northwest Slippersnails  (pdf)

Direct Feeding on Kelp by Littorina and Lacuna snails (Family Littorinidae) in the NE Pacific  (pdf)

Expanded Northern Range of Acanthodoris rhodoceras in Oregon  (pdf)

Freak Fissurellidea bimaculata (Dall, 1871) from Sooke, B.C. (pdf)

Hoof Snails (Family Hipponicidae) of B.C. and Washington  (pdf)

I.D. of a Boundary Bay, B.C. Opisthobranch  (pdf)

Japanese Nassa, Hima fratercula (Dunker, 1860) is more widely distributed than previously thought  (pdf)

Limpet Range Extension into BC  (pdf)

Mollusca Non-Grata:  The influence of top-down control and residence time on the abundance, distribution, and
behavior of non-native marine snails in Washington State  PhD dissertation of Emily W. Grason PhD  (pdf 8.29Mb)

Mollusks - Home Redecorating  (pdf)

Morphological, Behavioral and Reproductive Inducible Defenses in Two Species of Intertidal
Gastropods  Masters thesis of Christopher Brooks Morgan (pdf 1.1Mb)

Name correction for Odostomia on chiton featured in The Dredgings  (pdf)

New Records for the Distribution and a description of the egg capsules of the Japanese Nassa,
Hima fratercula (Dunker, 1860) Family Nassariidae in Ladysmith Harbour, B.C.  (pdf)

New Systematic Revisions Within the Vetigastropoda  (pdf)

A northern range extension for Oldroyd's Fragile Moon Snail, Calinaticina oldroydii (Dall, 1897)  (pdf)

The Nudibranchs of the Salish Sea  (pdf)

Of Corncobs and Flat Artichokes  (pdf) [Neptunea]

Olivella pycna Berry, 1935, least known of the Pacific Northwest Olives  (pdf)

Plesiocystiscus politulus, another California species found in British Columbia  (pdf)

Population Biology and Reproductive Ecology of Chlorostoma (Tegula) funebralis, An Intertidal Gastropod
Ph.D. dissertation of Erin Cooper Ph.D.  (pdf 2.56Mb)

The rare Elegant ribbed limpet, Scelidotoma bella (Gabb, 1865)  (pdf)

Records of the Red Abalone in British Columbia  (pdf)

The Secret Weapon of the Keyhole Limpet  (pdf)

Tidal migration patterns moderate thermal risk in the intertidal snail Nucella ostrina  PhD dissertation of Hilary Anne
Hayford PhD (pdf 3.74Mb

Update on Systematic Revisions within the Vetigastropoda  (pdf)

With Micro Molluscs Every Sample is a grab-bag!  (pdf)


An unexpected find of Carychium minimum Müller, 1774, an uncommon introduced land snail   (pdf)

Determining habitat suitability for the Rocky Mountain ridged mussel (Gonidea angulata, Lea 1838) in
Okanagan Lake, BC, Canada  (pdf)

Examining Fish Stomach Contents to Find a Rare Snail  (pdf)

Exotic species of land snails from Mount Vernon, Wa and Queen Elizabeth Park, Vancouver, BC (pdf)

Freshwater mollusks found during a survey for Potamopyrgus antipodarum  (pdf)

Invasion by Cernuella virgata in Tacoma (Update)  (pdf)

Land Snailing in Kitsap County  (pdf)

Marsh Snails in Boundary Bay  (pdf)

Modeling habitat suitability for the Rocky Mountain Ridged Mussel (Gonidea angulata), in Okanagan Lake,
British Columbia, Canada  Masters thesis of Roxanne May Snook (pdf 1.69Mb)

Monadenia Trapped Between Timbers  (pdf)

New species of arionid semi-slug found on Haida Gwaii [Staala gwaii]  (pdf)

On Finding a Species of Vertigo in Manchester State Park [WA]  (pdf)

Some Georgia Land Snails  (pdf)

Springs and the minute snails that inhabit them in the Puget Sound region  (pdf)

Terry Frest Honored with having a new species of freshwater snail named for him  (pdf)

U.S. Fish and Wildlife Service Rejects Fourteen Freshwater Snails of Twenty-six Pacific Northwest
Mollusks Proposed for Listing and sets a Dangerous Precedent  (pdf)

Zonitoides nitidus, one more new species find for me while walking my dog Zoe  (pdf)

     Corbicula fluminea:

The Asiatic Clam (Corbicula fluminea) found for first time in the Fraser River  (pdf)

The Asiatic Clam (Corbicula fluminea) 'Rediscovered' on Vancouver Island  (pdf)

The invasive Asian clam, Corbicula fluminea, found living in Lake Whatcom  (pdf)

Results of a visual survey of the lower Fraser River for the Asian Clam, Corbicula fluminea Mũller, 1774  (pdf)

     Potamopyrgus antipodarum:

A first report of the New Zealand mudsnail in British Columbia  (pdf)

Capitol Lake Update - New Zealand mud snail  (pdf)

A comparison of successful and unsuccessful invasive lineages of the New Zealand mud snail  (pdf)

New Zealand Mud Snails Found in Capitol Lake, Olympia, WA  (pdf)

Other Mollusks and Invertebrates

2nd Hand Snails via Hermit Crab By-catch - or - Let the Hermit Crabs do the work!  (pdf)

Chitoning the Salish Sea  (pdf)

Dentalium Encounters  (pdf)

The Harrowing Dilemma of a Female Pea Crab (pdf)

It Happened One Night  (pdf) [Loligo opalescens]

Molluscs Associated with a Long-Line West Coast Oyster Farm  (pdf)

A Parting Gift  (pdf) [sea star stomach contents]

Sea Star Catastrophe  (pdf)

Some observations of mass die-off of gumboot chitons  (pdf)

Spawning Season at Deception Pass  (pdf) [Mopalia swanii]

The Straight Hermit Crab  (pdf)

Field Trips/Shelling Adventures

A Micro Vacation on Roatan - 2013  (pdf)

An Icelandic Stopover - 2013  (pdf)

An Outing to Port Orchard - 2012  (pdf)

Collecting Freshwater Molluscs in Papua, New Guinea - 2009  (pdf)
Collecting Microshells from Weed Washings - 2014  (pdf)

Juan de Fuca Strait Field Trip - 2008  (pdf)

Marrowstone Point Field Trip - 2011  (pdf)

The Role of Serendipity in Shell Collecting  (pdf)

Shelling in Petersburg Alaska - 2009 (pdf 770k)

Slip Point Field Trip - 2010 (pdf)

Thetis Island Field Trip - 2012  (pdf 668k)

Two new species were found during a club field trip to Marrowstone Island - 2009 (pdf)

Victoria Field Trip - 2009 (pdf)
With Dr. Terry Frest on a mollusk survey and sampling trip to Idaho in 1990  (pdf)


A Focus Stacking Setup for Shell Photography  (pdf)

The Beginnings of the Pacific Northwest Shell Club  (pdf)  

Book Review - Spirals in time - The Secret Life and Curious Afterlife of Seashells  (pdf)

Happy 50th Birthday PNWSC (pdf)

In Memoriam - Roland Anderson  1946-2014  (pdf)

In Memoriam - Dr. Terrence J. Frest 2008  (pdf)

In Memoriam - Graham Jeffery 2014  (pdf)

In Memoriam - Dr. Ian McTaggart-Cowan 1910-2010  (pdf)

In Memoriam - Trevor Roberts  1914-2011 (pdf)

Name Changes of Recent Years  (html)

New Names for Some Pacific Northwest Locales  (pdf)

Tom Rice Awarded 2014 Neptunea Award by COA  (pdf)

Citation:  Pacific Northwest Shell Club,

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This page last revised:  11-20-2017

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